Drilliidae















































Drilliidae

Cymatosyrinx parciplicata 003.jpg
An apertural view of a shell of Cymatosyrinx parciplicata

Scientific classification edit
Kingdom:
Animalia
Phylum:
Mollusca
Class:
Gastropoda

Clade:

Caenogastropoda

Clade:

Hypsogastropoda

Clade:

Neogastropoda
Superfamily:
Conoidea
Family:
Drilliidae
Olsson, 1964
Genera

See text



Synonyms[1]

Clavidae Casey, 1904 (non McCrady, 1859: preoccupied)



The Drilliidae are a taxonomic family of small predatory sea snails with high-spired shells. They are classified as marine gastropod mollusks in the superfamily Conoidea.[2]


This family has no subfamilies.




Contents






  • 1 Description


  • 2 Taxonomic history


  • 3 Taxonomy


  • 4 References


  • 5 External links





Description


The shell is claviform with a tall spire, or squatly conical to biconical. In many species, the siphonal canal is truncated. The aperture is U-shaped with a parietal callus pad.


The sculpture of the shell shows prominent axial ribs with a polished surface. Most species have a dorsal varix (transverse elevation), except in the genera Cymatosyrinx, Elaeocyma and Splendrillia. The protoconch can be smooth or very carinate. The ovate operculum has a terminal nucleus.[3]


The radula of the species in this family have characteristically five teeth in each row (formula : 1 + 1 + 1 + 1 + 1) with a vestigial central tooth, comb-like lateral teeth and a pair of flat-pointed, slender marginal teeth.[4]


The foregut structure of the anterior alimentary system has the least derived foregut anatomy of all the families in the superfamily Conoidea.[5]



Taxonomic history


The genera in this family were originally separated from the subfamily Clavitulinae and classified by H.& A. Adams (1858) under subfamily Turritinae, because their operculum has a terminal instead of a central nucleus. In 1942, the species with a U-shaped sinus and a parietal callus pad were brought by Powell in the new subfamily Clavinae. In 1966, Morrison proposed the subfamily Drillinae ( = Clavinae) for the species with a stenoglossan radula with comb-like lateral teeth. Finally, in 1993, Taylor et al. proposed the promotion of Drillinae from subfamily to the family level Drilliidae.[3][6]



Taxonomy


Genera in the family Drilliidae include:[1]





  • Acinodrillia Kilburn, 1988


  • Agladrillia Woodring, 1928


  • Bellaspira Conrad, 1868


  • Calliclava McLean, 1971


  • Cerodrillia Bartsch & Rehder, 1939


  • Clathrodrillia Dall, 1918


  • Clavus Montfort, 1810


  • Conopleura Hinds, 1844


  • Crassopleura Monterosato, 1884


  • Cruziturricula Marks, 1951


  • Cymatosyrinx Dall, 1889


  • Decoradrillia Fallon, 2016


  • Douglassia Bartsch, 1934


  • Drillia Gray, 1838


  • Elaeocyma Dall, 1918


  • Eumetadrillia Woodring, 1928


  • Fenimorea Bartsch, 1934


  • Fusiturricula Woodring, 1928


  • Globidrillia Woodring, 1928


  • Hauturua Powell, 1942


  • Imaclava Bartsch, 1944


  • Iredalea Oliver, 1915


  • Kylix Dall, 1919


  • Leptadrillia Woodring, 1928


  • Lissodrillia Bartsch & Rehder, 1943


  • Neodrillia Bartsch, 1943


  • Orrmaesia Kilburn, 1988


  • Paracuneus Laseron, 1954


  • Plagiostropha Melvill, 1927

  • Pleurofusia de Gregorio, 1890


  • Sedilia Fargo, 1953


  • Spirotropis Sars, 1878


  • Splendrillia Hedley, 1922


  • Stenodrillia Korobkov, 1955


  • Syntomodrillia Hedley, 1922


  • Wairarapa Vella, 1954




Genera brought into synonymy





  • Brephodrillia Pilsbry & Lowe, 1932: synonym of Iredalea


  • Clavicantha Swainson, 1840: synonym of Clavus Montfort, 1810


  • Eldridgea Bartsch, 1934: synonym of Clavus Montfort, 1810


  • Fusisyrinx Bartsch, 1934: synonym of Fusiturricula Woodring, 1928


  • Tylotia Melvill, 1917: synonym of Clavus Montfort, 1810


  • Tylotiella Habe, 1958: synonym of Clavus Montfort, 1810



Genera moved to other families





  • Austroclavus (unassigned to a family within Conoidea)


  • Brachytoma moved to the family Pseudomelatomidae


  • Sediliopsis Petuch, 1988 : moved to the family Pseudomelatomidae



References





  1. ^ ab WoRMS (2010). Drilliidae. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=23032 on1 September 2011


  2. ^ Bouchet, P.; Rocroi, J.-P. (2005). "Classification and Nomenclator of Gastropod Families". Malacologia. 47 (1–2)..mw-parser-output cite.citation{font-style:inherit}.mw-parser-output q{quotes:"""""""'""'"}.mw-parser-output code.cs1-code{color:inherit;background:inherit;border:inherit;padding:inherit}.mw-parser-output .cs1-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/6/65/Lock-green.svg/9px-Lock-green.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .cs1-lock-limited a,.mw-parser-output .cs1-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Lock-gray-alt-2.svg/9px-Lock-gray-alt-2.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .cs1-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/a/aa/Lock-red-alt-2.svg/9px-Lock-red-alt-2.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registration{color:#555}.mw-parser-output .cs1-subscription span,.mw-parser-output .cs1-registration span{border-bottom:1px dotted;cursor:help}.mw-parser-output .cs1-hidden-error{display:none;font-size:100%}.mw-parser-output .cs1-visible-error{font-size:100%}.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registration,.mw-parser-output .cs1-format{font-size:95%}.mw-parser-output .cs1-kern-left,.mw-parser-output .cs1-kern-wl-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right,.mw-parser-output .cs1-kern-wl-right{padding-right:0.2em}


  3. ^ ab Chen-Kwoh Chang, Small Turrids of Taiwan, Chapter 3 , History and Taxonomy of the Clavidae; June 1, 2001[permanent dead link]


  4. ^ Kantor, Yuri I; John D.Taylor (2000). "Formation of marginal radular teeth in Conoidea (Neogastropoda) and the evolution of the hypodermic envenomation mechanism". Journal of Zoology. Cambridge University Press. 252 (2): 251–262. doi:10.1111/j.1469-7998.2000.tb00620.x.


  5. ^ Brian Morton, The Malacofauna of Hong Kong and southern China


  6. ^ Taylor, J.D., Kantor, Y.I& Sysoev, A.V.,1993 . Foregut anatomy, feeding mechanisms, relationships and classification of the Conoidea (=Toxoglossas) (Gastropoda) Bull. Nat. Hist. Mus. Lond .(Zool) 5(2): 125-170



  • The Taxonomicon


External links



  • PHILLIP J. Jr. FALLON, Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species; Zootaxa, [S.l.], v. 4090, n. 1, p. 1–363, mar. 2016. ISSN 1175-5334

  • WMSDB - Worldwide Mollusc Species Data Base: family Drilliidae

















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